/* =========================
   DESIGN SYSTEM
========================= */

:root {
    --background: #f3f4f0;
    --surface: #ffffff;
    --surface-soft: #e8ece7;

    --text-primary: #16202a;
    --text-secondary: #59646d;

    --accent: #315c59;
    --accent-dark: #234440;
    --accent-soft: #dce7e3;

    --dark: #101a1d;

    --publication-accent: #a7cec4;
    --publication-accent-light: #b8ddd3;

    --border: rgba(22, 32, 42, 0.13);

    --content-width: 1240px;

    --radius-small: 10px;
    --radius-medium: 20px;
    --radius-large: 32px;

    --shadow:
        0 20px 50px
        rgba(21, 36, 38, 0.08);
}


/* =========================
   RESET
========================= */

* {
    box-sizing: border-box;
}

html {
    scroll-behavior: smooth;
}

body {
    margin: 0;
    overflow-x: hidden;

    font-family:
        -apple-system,
        BlinkMacSystemFont,
        "Segoe UI",
        Helvetica,
        Arial,
        sans-serif;

    background: var(--background);
    color: var(--text-primary);
    line-height: 1.6;

    -webkit-font-smoothing: antialiased;
}

body.menu-open {
    overflow: hidden;
}

a {
    color: inherit;
    text-decoration: none;
}

button,
input,
textarea {
    font: inherit;
}

img {
    display: block;
    max-width: 100%;
}

p {
    margin-top: 0;
}

section {
    scroll-margin-top: 90px;
}


/* =========================
   ACCESSIBILITY
========================= */

a:focus-visible,
button:focus-visible,
input:focus-visible,
textarea:focus-visible {
    outline: 2px solid var(--accent);
    outline-offset: 4px;
}


/* =========================
   SKIP LINK
========================= */

.skip-link {
    position: fixed;
    top: -100px;
    left: 20px;
    z-index: 9999;

    padding: 12px 18px;

    border-radius: 8px;
    background: var(--dark);
    color: #ffffff;
}

.skip-link:focus {
    top: 20px;
}


/* =========================
   NAVIGATION
========================= */

.site-header {
    position: sticky;
    top: 0;
    z-index: 1000;

    border-bottom: 1px solid rgba(22, 32, 42, 0.08);

    background: rgba(243, 244, 240, 0.90);

    backdrop-filter: blur(18px);
    -webkit-backdrop-filter: blur(18px);
}

.nav-container {
    width: min(92%, var(--content-width));
    height: 78px;
    margin: 0 auto;

    display: flex;
    align-items: center;
    justify-content: space-between;

    gap: 36px;
}

.brand {
    width: 44px;
    height: 44px;
    flex: 0 0 auto;

    display: grid;
    place-items: center;

    border-radius: 50%;

    background: var(--text-primary);
    color: #ffffff;

    font-size: 0.86rem;
    font-weight: 700;
    letter-spacing: 0.08em;

    transition:
        background 180ms ease,
        transform 180ms ease;
}

.brand:hover {
    background: var(--accent);
    transform: translateY(-1px);
}

.site-nav {
    display: flex;
    align-items: center;

    gap: clamp(17px, 2vw, 31px);

    font-size: 0.83rem;
    font-weight: 550;
}

.site-nav a {
    position: relative;

    color: var(--text-secondary);
    white-space: nowrap;

    transition: color 180ms ease;
}

.site-nav a::after {
    content: "";

    position: absolute;
    left: 0;
    bottom: -7px;

    width: 100%;
    height: 1px;

    background: var(--text-primary);

    transform: scaleX(0);
    transform-origin: right;

    transition: transform 180ms ease;
}

.site-nav a:hover {
    color: var(--text-primary);
}

.site-nav a:hover::after {
    transform: scaleX(1);
    transform-origin: left;
}

.menu-toggle {
    display: none;

    width: 46px;
    height: 46px;

    padding: 0;
    border: 0;

    background: transparent;
    cursor: pointer;
}

.menu-toggle span {
    display: block;

    width: 22px;
    height: 1.5px;

    margin: 6px auto;

    background: var(--text-primary);

    transition: transform 180ms ease;
}


/* =========================
   HERO
========================= */

.hero {
    width: min(92%, var(--content-width));
    min-height: calc(100svh - 78px);
    margin: 0 auto;

    padding:
        clamp(72px, 8vw, 110px)
        0
        clamp(90px, 9vw, 125px);

    display: flex;
    align-items: center;
}

.hero-grid {
    width: 100%;

    display: grid;

    grid-template-columns:
        minmax(0, 1.2fr)
        minmax(340px, 0.8fr);

    gap: clamp(44px, 6vw, 90px);

    align-items: center;
}

.hero-content {
    min-width: 0;
    position: relative;
    z-index: 2;
}

.eyebrow {
    margin-bottom: 24px;

    color: var(--accent);

    font-size: 0.78rem;
    font-weight: 700;
    letter-spacing: 0.16em;

    text-transform: uppercase;
}

.hero h1 {
    max-width: 850px;

    margin: 0 0 34px;

    font-size: clamp(3.7rem, 6.6vw, 6.9rem);

    font-weight: 650;
    letter-spacing: -0.06em;
    line-height: 0.92;
}

.hero h1 span {
    color: var(--accent);
    white-space: nowrap;
}

.hero-statement {
    max-width: 760px;
    margin-bottom: 34px;

    color: var(--text-secondary);

    font-size: clamp(1.28rem, 2.1vw, 1.9rem);

    line-height: 1.42;
    letter-spacing: -0.025em;
}

.research-tags {
    max-width: 760px;

    display: flex;
    flex-wrap: wrap;

    gap: 9px;
    margin-bottom: 42px;
}

.research-tags span {
    display: inline-flex;
    align-items: center;

    min-height: 38px;

    padding: 7px 13px;

    border: 1px solid var(--border);
    border-radius: 50px;

    background: rgba(255, 255, 255, 0.45);
    color: var(--text-secondary);

    font-size: 0.78rem;
}

.hero-actions {
    max-width: 760px;

    display: grid;

    grid-template-columns:
        max-content
        max-content;

    justify-content: space-between;
    align-items: center;

    gap: 18px;
}

.button {
    display: inline-flex;
    align-items: center;
    justify-content: center;

    min-height: 50px;

    padding: 0 23px;

    border-radius: 50px;

    font-size: 0.88rem;
    font-weight: 650;

    transition:
        transform 180ms ease,
        background 180ms ease,
        color 180ms ease,
        border-color 180ms ease;
}

.button:hover {
    transform: translateY(-2px);
}

.button-primary {
    background: var(--text-primary);
    color: #ffffff;
}

.button-primary:hover {
    background: var(--accent);
}

.button-secondary {
    border: 1px solid var(--border);

    background: rgba(255, 255, 255, 0.52);
    color: var(--text-primary);
}

.button-secondary:hover {
    border-color: var(--accent);
    background: rgba(255, 255, 255, 0.9);
}


/* =========================
   HERO ARTWORK
========================= */

.hero-media {
    position: relative;

    min-width: 0;
    min-height: 620px;

    margin: 0;

    display: grid;
    place-items: center;

    overflow: visible;

    background: transparent;
    border: 0;
    border-radius: 0;
    box-shadow: none;
}

.hero-artwork {
    width: auto;

    height: min(64vh, 620px);

    max-width: none;
    max-height: 620px;

    margin: 0 -30px 0 auto;

    object-fit: contain;

    filter:
        drop-shadow(
            0 26px 35px
            rgba(49, 92, 89, 0.08)
        );
}


/* =========================
   SHARED SECTIONS
========================= */

.section {
    width: min(92%, var(--content-width));
    margin: 0 auto;

    padding: 130px 0;
}

.section-alt {
    width: 100%;
    max-width: none;

    padding:
        130px
        max(
            4%,
            calc((100vw - var(--content-width)) / 2)
        );

    background: var(--surface-soft);
}

.section-grid {
    display: grid;

    grid-template-columns:
        0.65fr
        1.35fr;

    gap: clamp(50px, 10vw, 150px);
}

.section-heading h2,
.section-header h2,
.communication-grid h2 {
    margin: 0;

    font-size: clamp(2.8rem, 5vw, 5.4rem);

    font-weight: 600;
    letter-spacing: -0.055em;
    line-height: 0.96;
}

.section-content {
    max-width: 760px;
}

.section-content > p {
    color: var(--text-secondary);

    font-size: 1.02rem;
    line-height: 1.85;
}

.large-copy {
    margin-bottom: 36px !important;

    color: var(--text-primary) !important;

    font-size: clamp(1.7rem, 3vw, 2.65rem) !important;

    line-height: 1.35 !important;
    letter-spacing: -0.035em;
}

.section-header {
    display: flex;
    align-items: flex-end;
    justify-content: space-between;

    gap: 60px;
    margin-bottom: 70px;
}

.section-header > p {
    max-width: 430px;
    margin-bottom: 0;

    color: var(--text-secondary);
    font-size: 0.95rem;
}


/* =========================
   ABOUT
========================= */

.about-grid {
    grid-template-columns:
        minmax(300px, 0.8fr)
        minmax(0, 1.2fr);

    align-items: start;
}

.about-left {
    display: flex;
    flex-direction: column;
    align-items: flex-start;
}

.about-left .section-heading {
    width: 100%;
}

.about-portrait {
    width: 100%;
    max-width: 430px;

    margin: 55px 0 0;

    overflow: hidden;

    border-radius: var(--radius-medium);

    background: var(--surface-soft);

    box-shadow: var(--shadow);
}

.about-portrait img {
    width: 100%;

    aspect-ratio: 3 / 4;

    object-fit: cover;
    object-position: center;

    transition: transform 300ms ease;
}

.about-portrait:hover img {
    transform: scale(1.02);
}

.about-copy {
    min-width: 0;
}

.about-copy > p {
    color: var(--text-secondary);

    font-size: 1.02rem;
    line-height: 1.85;
}


/* =========================
   DOCTORAL CREDENTIAL
========================= */

.doctoral-credential {
    margin: 0 0 38px;

    padding: 22px 24px;

    display: flex;
    flex-direction: column;
    align-items: flex-start;

    gap: 6px;

    border-left: 3px solid var(--accent);

    border-radius:
        0
        var(--radius-small)
        var(--radius-small)
        0;

    background: var(--accent-soft);
}

.doctoral-credential-label {
    color: var(--accent);

    font-size: 0.70rem;
    font-weight: 750;
    letter-spacing: 0.12em;

    text-transform: uppercase;
}

.doctoral-credential strong {
    color: var(--text-primary);

    font-size: 1rem;
    font-weight: 700;
    line-height: 1.45;
}

.doctoral-credential-meta {
    color: var(--text-secondary);

    font-size: 0.80rem;
    line-height: 1.5;
}


/* =========================
   RESEARCH CARDS
========================= */

.research-grid {
    display: grid;

    grid-template-columns:
        repeat(2, minmax(0, 1fr));

    gap: 14px;
}

.research-card {
    min-height: 320px;

    display: flex;
    flex-direction: column;
    justify-content: flex-end;

    padding: clamp(28px, 4vw, 48px);

    border: 1px solid rgba(22, 32, 42, 0.08);
    border-radius: var(--radius-medium);

    background: rgba(255, 255, 255, 0.58);

    transition:
        transform 180ms ease,
        background 180ms ease,
        box-shadow 180ms ease;
}

.research-card:hover {
    transform: translateY(-4px);

    background: rgba(255, 255, 255, 0.88);

    box-shadow:
        0 15px 35px
        rgba(21, 36, 38, 0.05);
}

.research-card h3 {
    max-width: 430px;

    margin: 0 0 18px;

    font-size: clamp(1.5rem, 2.4vw, 2.25rem);

    font-weight: 600;
    line-height: 1.12;
    letter-spacing: -0.035em;
}

.research-card p {
    max-width: 500px;
    margin-bottom: 0;

    color: var(--text-secondary);

    font-size: 0.94rem;
    line-height: 1.75;
}


/* =========================
   FEATURED RESEARCH VISUAL
========================= */

.research-feature {
    margin: 90px 0 0;

    padding-top: 55px;

    border-top: 1px solid var(--border);
}

.research-feature-header {
    width: 100%;
    max-width: 900px;

    margin: 0 auto 42px;

    text-align: center;
}

.research-feature-header .eyebrow {
    margin-bottom: 18px;
}

.research-feature-header h3 {
    max-width: 850px;

    margin: 0 auto;

    font-size: clamp(2rem, 4vw, 4rem);

    font-weight: 550;
    letter-spacing: -0.045em;
    line-height: 1.08;
}

.research-feature-header > p {
    max-width: 760px;

    margin: 24px auto 0;

    color: var(--text-secondary);

    font-size: 0.95rem;
    line-height: 1.75;
}

.research-feature-image {
    display: block;
    width: 100%;

    overflow: hidden;

    border: 1px solid var(--border);
    border-radius: var(--radius-medium);

    background: #ffffff;

    box-shadow: var(--shadow);

    transition:
        transform 180ms ease,
        box-shadow 180ms ease;
}

.research-feature-image:hover {
    transform: translateY(-3px);

    box-shadow:
        0 24px 55px
        rgba(21, 36, 38, 0.12);
}

.research-feature-image img {
    width: 100%;
    height: auto;
}

.research-feature figcaption {
    display: flex;
    align-items: center;
    justify-content: center;
    flex-wrap: wrap;

    gap: 8px;
    margin-top: 16px;

    text-align: center;

    color: var(--text-secondary);

    font-size: 0.75rem;
    line-height: 1.5;
}

.research-feature figcaption span {
    color: var(--text-primary);
    font-weight: 650;
}

.research-feature figcaption span::after {
    content: "·";

    margin-left: 8px;

    color: var(--text-secondary);
    font-weight: 400;
}

.research-feature figcaption a {
    color: var(--accent);
    font-weight: 650;

    transition: color 180ms ease;
}

.research-feature figcaption a:hover {
    color: var(--accent-dark);
}


/* =========================
   RESEARCH EXPERIENCE
========================= */

.timeline {
    border-top: 1px solid var(--border);
}

.timeline-item {
    display: grid;

    grid-template-columns:
        0.55fr
        1.45fr;

    gap: 60px;

    padding: 42px 0;

    border-bottom: 1px solid var(--border);
}

.timeline-date {
    color: var(--accent);

    font-size: 0.79rem;
    font-weight: 700;
    letter-spacing: 0.08em;

    text-transform: uppercase;
}

.timeline-content h3 {
    margin: 0 0 8px;

    font-size: clamp(1.4rem, 2.3vw, 2rem);

    font-weight: 600;
    letter-spacing: -0.025em;
}

.institution {
    margin-bottom: 18px;

    color: var(--text-primary) !important;
    font-weight: 600;
}

.timeline-content p {
    max-width: 760px;

    color: var(--text-secondary);
    line-height: 1.75;
}


/* =========================
   DARK SECTION
========================= */

.section-dark {
    width: 100%;
    max-width: none;

    padding:
        130px
        max(
            4%,
            calc((100vw - var(--content-width)) / 2)
        );

    background: var(--dark);
    color: #ffffff;
}

.section-dark .section-header > p {
    color: rgba(255, 255, 255, 0.58);
}


/* =========================
   PUBLICATIONS
========================= */

.publications-list {
    border-top: 1px solid rgba(255, 255, 255, 0.18);
}

.publication-item {
    display: grid;

    grid-template-columns:
        minmax(150px, 0.35fr)
        minmax(0, 1.65fr);

    gap: clamp(35px, 7vw, 100px);

    padding: 52px 0;

    border-bottom: 1px solid rgba(255, 255, 255, 0.14);
}

.publication-meta {
    display: flex;
    flex-direction: column;
    align-items: flex-start;

    gap: 12px;
}

.publication-year {
    color: #ffffff;

    font-size: 0.82rem;
    font-weight: 700;
    letter-spacing: 0.08em;
}

.publication-type,
.publication-status {
    display: inline-flex;
    align-items: center;

    padding: 6px 10px;

    border: 1px solid rgba(255, 255, 255, 0.2);
    border-radius: 30px;

    color: rgba(255, 255, 255, 0.65);

    font-size: 0.68rem;
    font-weight: 650;
    line-height: 1.25;
    letter-spacing: 0.06em;

    text-transform: uppercase;
}

.publication-status {
    border-color: rgba(167, 206, 196, 0.55);

    color: var(--publication-accent-light);

    background: rgba(167, 206, 196, 0.08);
}

.publication-content {
    max-width: 900px;
}

.publication-authors {
    margin-bottom: 17px;

    color: rgba(255, 255, 255, 0.61);

    font-size: 0.87rem;
    line-height: 1.65;
}

.publication-authors strong {
    color: var(--publication-accent);
    font-weight: 700;
}

.publication-content h3 {
    max-width: 880px;

    margin: 0 0 22px;

    color: #ffffff;

    font-size: clamp(1.55rem, 3vw, 2.65rem);

    font-weight: 550;
    letter-spacing: -0.035em;
    line-height: 1.18;
}

.publication-journal {
    margin-bottom: 20px;

    color: rgba(255, 255, 255, 0.65);

    font-size: 0.9rem;
    line-height: 1.65;
}

.publication-note {
    margin-bottom: 0;

    color: var(--publication-accent-light);

    font-size: 0.78rem;
    font-weight: 600;
    letter-spacing: 0.03em;
}

.publications-footer {
    display: flex;
    justify-content: flex-end;

    margin-top: 46px;
}

.publication-record-link {
    display: inline-flex;
    align-items: center;
    justify-content: center;

    gap: 10px;

    min-height: 48px;

    padding: 0 20px;

    border: 1px solid var(--publication-accent);
    border-radius: 50px;

    background: var(--publication-accent);
    color: var(--dark);

    font-size: 0.8rem;
    font-weight: 750;

    transition:
        background 180ms ease,
        border-color 180ms ease,
        transform 180ms ease,
        box-shadow 180ms ease;
}

.publication-record-link:hover {
    background: var(--publication-accent-light);
    border-color: var(--publication-accent-light);

    transform: translateY(-2px);

    box-shadow:
        0 10px 26px
        rgba(0, 0, 0, 0.18);
}

.publication-link {
    display: inline-flex;
    align-items: center;
    justify-content: center;

    gap: 9px;

    min-height: 42px;
    margin-top: 4px;

    padding: 9px 16px;

    border: 1px solid var(--publication-accent);
    border-radius: 50px;

    background: var(--publication-accent);
    color: var(--dark);

    font-size: 0.8rem;
    font-weight: 750;
    letter-spacing: 0.015em;

    box-shadow:
        0 7px 20px
        rgba(0, 0, 0, 0.12);

    transition:
        background 180ms ease,
        color 180ms ease,
        border-color 180ms ease,
        transform 180ms ease,
        box-shadow 180ms ease;
}

.publication-link:hover {
    background: #c4e2da;
    border-color: #c4e2da;
    color: #0c1719;

    transform: translateY(-2px);

    box-shadow:
        0 10px 26px
        rgba(0, 0, 0, 0.18);
}


/* =========================
   EXPERTISE
========================= */

.expertise-grid {
    display: grid;

    grid-template-columns:
        repeat(4, minmax(0, 1fr));

    border-top: 1px solid var(--border);
    border-bottom: 1px solid var(--border);
}

.expertise-grid article {
    padding: 38px 30px;

    border-right: 1px solid var(--border);
}

.expertise-grid article:first-child {
    padding-left: 0;
}

.expertise-grid article:last-child {
    padding-right: 0;
    border-right: 0;
}

.expertise-grid h3 {
    min-height: 60px;

    margin: 0 0 28px;

    font-size: 1.15rem;
    font-weight: 650;
}

.expertise-grid ul {
    margin: 0;
    padding: 0;

    list-style: none;
}

.expertise-grid li {
    padding: 8px 0;

    color: var(--text-secondary);
    font-size: 0.9rem;
}


/* =========================
   SCIENCE COMMUNICATION
========================= */

.communication-section {
    width: 100%;
    max-width: none;

    padding:
        130px
        max(
            4%,
            calc((100vw - var(--content-width)) / 2)
        );

    background: var(--accent-soft);
}

.communication-grid {
    display: grid;

    grid-template-columns:
        0.8fr
        1.2fr;

    gap: clamp(50px, 10vw, 150px);
}

.communication-feature {
    max-width: 720px;
}

.communication-feature h3 {
    margin: 0 0 30px;

    font-size: clamp(2rem, 4vw, 4.3rem);

    font-weight: 550;
    letter-spacing: -0.045em;
    line-height: 1.08;
}

.communication-feature > p:not(.eyebrow) {
    max-width: 620px;

    color: var(--text-secondary);

    font-size: 1rem;
    line-height: 1.8;
}

.communication-meta {
    display: flex;
    flex-wrap: wrap;

    gap: 9px;
    margin-top: 34px;
}

.communication-badge {
    display: inline-flex;
    align-items: center;
    justify-content: center;

    padding: 8px 14px;

    border: 1px solid rgba(49, 92, 89, 0.30);
    border-radius: 50px;

    background: rgba(255, 255, 255, 0.55);
    color: var(--accent-dark);

    font-size: 0.74rem;
    font-weight: 700;
    line-height: 1.3;

    white-space: nowrap;
}


/* =========================
   GENE & TONIC FEATURE PHOTOS
========================= */

.communication-feature-gallery {
    width: 100%;
    max-width: 1080px;

    margin: 64px auto 0;

    display: grid;

    grid-template-columns:
        repeat(2, minmax(0, 1fr));

    gap: 16px;
}

.communication-feature-photo {
    position: relative;

    margin: 0;

    overflow: hidden;

    aspect-ratio: 16 / 10;

    border-radius: var(--radius-medium);

    background: rgba(255, 255, 255, 0.55);

    box-shadow: var(--shadow);
}

.communication-feature-photo img {
    width: 100%;
    height: 100%;

    object-fit: cover;

    pointer-events: none;

    user-select: none;
    -webkit-user-select: none;

    -webkit-user-drag: none;
    -webkit-touch-callout: none;

    transition: transform 300ms ease;
}

.communication-feature-photo:hover img {
    transform: scale(1.015);
}


/* =========================
   GENE & TONIC EVENT MOSAIC
========================= */

.communication-mosaic {
    width: 100%;
    max-width: 1080px;

    margin: 14px auto 0;

    display: grid;

    grid-template-columns:
        repeat(4, minmax(0, 1fr));

    gap: 10px;
}

.communication-mosaic-item {
    position: relative;

    margin: 0;

    overflow: hidden;

    aspect-ratio: 4 / 3;

    border-radius: 12px;

    background: rgba(255, 255, 255, 0.55);
}

.communication-mosaic-item img {
    width: 100%;
    height: 100%;

    object-fit: cover;

    pointer-events: none;

    user-select: none;
    -webkit-user-select: none;

    -webkit-user-drag: none;
    -webkit-touch-callout: none;

    transition: transform 250ms ease;
}

.communication-mosaic-item:hover img {
    transform: scale(1.02);
}


/* Transparent interaction layer.
   This discourages casual long-press / drag / image interaction. */

.communication-feature-photo::after,
.communication-mosaic-item::after {
    content: "";

    position: absolute;
    inset: 0;
    z-index: 5;

    background: transparent;

    user-select: none;
    -webkit-user-select: none;
    -webkit-touch-callout: none;
}

.communication-gallery-note {
    margin: 28px 0 0;

    color: var(--accent-dark);

    font-size: 0.74rem;
    font-weight: 700;
    letter-spacing: 0.09em;

    text-align: center;
    text-transform: uppercase;
}


/* =========================
   ACTIVITIES
========================= */

.activity-list {
    margin: 50px 0 30px;

    border-top: 1px solid var(--border);
}

.activity-list div {
    display: grid;

    grid-template-columns:
        0.6fr
        1.4fr;

    gap: 30px;

    padding: 22px 0;

    border-bottom: 1px solid var(--border);
}

.activity-list span {
    color: var(--text-secondary);
    font-size: 0.8rem;
}

.activity-list strong {
    font-size: 0.95rem;
    font-weight: 600;
}

.cv-note {
    margin-top: 32px;

    font-size: 0.82rem !important;
}


/* =========================
   CONTACT
========================= */

.contact-section {
    padding: 140px 4%;

    background: var(--text-primary);
    color: #ffffff;
}

.contact-inner {
    width: min(100%, var(--content-width));
    margin: 0 auto;
}

.contact-header {
    max-width: 1080px;
    margin-bottom: 70px;
}

.contact-header h2 {
    max-width: 1050px;

    margin: 0 0 32px;

    font-size: clamp(3rem, 7vw, 7rem);

    font-weight: 550;
    letter-spacing: -0.06em;
    line-height: 0.98;
}

.contact-intro {
    max-width: 680px;
    margin: 0;

    color: rgba(255, 255, 255, 0.64);

    font-size: 1rem;
    line-height: 1.8;
}

.contact-layout {
    display: grid;

    grid-template-columns:
        minmax(0, 1.45fr)
        minmax(250px, 0.55fr);

    gap: clamp(60px, 8vw, 120px);

    align-items: start;
}

.contact-form {
    width: 100%;
    max-width: 820px;
}

.form-row {
    display: grid;

    grid-template-columns:
        repeat(2, minmax(0, 1fr));

    gap: 18px;
}

.form-field {
    width: 100%;
    margin-bottom: 24px;
}

.form-field label {
    display: flex;
    align-items: center;
    justify-content: space-between;

    gap: 12px;
    margin-bottom: 9px;

    color: rgba(255, 255, 255, 0.8);

    font-size: 0.77rem;
    font-weight: 650;
    letter-spacing: 0.03em;
}

.form-field label span {
    color: rgba(255, 255, 255, 0.38);

    font-size: 0.68rem;
    font-weight: 500;
}

.form-field input,
.form-field textarea {
    display: block;
    width: 100%;

    margin: 0;

    border: 1px solid rgba(255, 255, 255, 0.18);
    border-radius: 14px;
    outline: none;

    appearance: none;
    -webkit-appearance: none;

    background: rgba(255, 255, 255, 0.055);
    color: #ffffff;

    font-size: 0.95rem;
    box-shadow: none;

    transition:
        border-color 180ms ease,
        background 180ms ease,
        box-shadow 180ms ease;
}

.form-field input {
    height: 56px;

    padding: 0 16px;
}

.form-field textarea {
    min-height: 190px;

    padding: 15px 16px;

    line-height: 1.6;
    resize: vertical;
}

.form-field input:hover,
.form-field textarea:hover {
    border-color: rgba(255, 255, 255, 0.32);
}

.form-field input:focus,
.form-field textarea:focus {
    border-color: var(--publication-accent);

    background: rgba(255, 255, 255, 0.08);

    box-shadow:
        0 0 0 3px
        rgba(167, 206, 196, 0.10);
}

.form-status {
    min-height: 24px;
    margin-top: 18px;

    font-size: 0.8rem;
    line-height: 1.6;
}

.form-status.success {
    color: var(--publication-accent-light);
}

.form-status.error {
    color: #f0b6b6;
}

.form-submit:disabled {
    cursor: wait;
    opacity: 0.72;

    transform: none;
    box-shadow: none;
}

.form-honeypot {
    position: absolute !important;

    width: 1px !important;
    height: 1px !important;

    margin: -1px !important;
    padding: 0 !important;

    overflow: hidden !important;

    clip: rect(0, 0, 0, 0) !important;
    clip-path: inset(50%) !important;

    border: 0 !important;

    white-space: nowrap !important;
}

.form-submit {
    display: inline-flex;
    align-items: center;
    justify-content: center;

    gap: 10px;

    min-height: 50px;

    padding: 0 21px;

    border: 1px solid var(--publication-accent);
    border-radius: 50px;

    appearance: none;
    -webkit-appearance: none;

    background: var(--publication-accent);
    color: var(--dark);

    cursor: pointer;

    font-size: 0.84rem;
    font-weight: 750;

    box-shadow:
        0 8px 24px
        rgba(0, 0, 0, 0.14);

    transition:
        background 180ms ease,
        border-color 180ms ease,
        transform 180ms ease,
        box-shadow 180ms ease;
}

.form-submit:hover {
    background: #c4e2da;
    border-color: #c4e2da;

    transform: translateY(-2px);

    box-shadow:
        0 11px 28px
        rgba(0, 0, 0, 0.20);
}

.contact-profiles {
    width: 100%;
    padding-top: 2px;
}

.contact-profiles-label {
    margin: 0 0 22px;

    color: rgba(255, 255, 255, 0.42);

    font-size: 0.7rem;
    font-weight: 700;
    letter-spacing: 0.14em;

    text-transform: uppercase;
}

.contact-profiles > a {
    display: flex;
    align-items: center;
    justify-content: space-between;

    gap: 20px;

    width: 100%;

    padding: 18px 0;

    border-bottom: 1px solid rgba(255, 255, 255, 0.14);

    color: rgba(255, 255, 255, 0.88);

    font-size: 0.95rem;
    font-weight: 600;

    transition:
        color 180ms ease,
        padding-left 180ms ease;
}

.contact-profiles > a:first-of-type {
    border-top: 1px solid rgba(255, 255, 255, 0.14);
}

.contact-profiles > a:hover {
    padding-left: 5px;

    color: var(--publication-accent-light);
}

.contact-profiles > a span:last-child {
    color: var(--publication-accent);
}

.contact-privacy {
    max-width: 320px;

    margin: 28px 0 0;

    color: rgba(255, 255, 255, 0.40);

    font-size: 0.72rem;
    line-height: 1.7;
}


/* =========================
   FOOTER
========================= */

.site-footer {
    min-height: 110px;

    display: flex;
    align-items: center;
    justify-content: space-between;

    gap: 30px;

    padding:
        0
        max(
            4%,
            calc((100vw - var(--content-width)) / 2)
        );

    border-top: 1px solid rgba(255, 255, 255, 0.1);

    background: var(--dark);

    color: rgba(255, 255, 255, 0.62);

    font-size: 0.78rem;
}

.footer-identity {
    display: flex;
    align-items: center;

    gap: 14px;
}

.footer-identity strong {
    display: grid;
    place-items: center;

    width: 34px;
    height: 34px;

    border: 1px solid rgba(255, 255, 255, 0.22);
    border-radius: 50%;

    color: #ffffff;
    font-size: 0.65rem;
}

.footer-links {
    display: flex;
    align-items: center;
    justify-content: center;
    flex-wrap: wrap;

    gap: 20px;
}

.footer-links a {
    color: rgba(255, 255, 255, 0.62);

    font-size: 0.76rem;
    font-weight: 600;

    transition: color 180ms ease;
}

.footer-links a:hover {
    color: var(--publication-accent-light);
}

.footer-label {
    margin: 0;

    color: rgba(255, 255, 255, 0.5);
}


/* =========================
   TABLET / IPAD
========================= */

@media (max-width: 1100px) {

    .site-nav {
        gap: 16px;
        font-size: 0.75rem;
    }

    .site-nav a:nth-child(6) {
        display: none;
    }

    .hero-grid {
        grid-template-columns:
            minmax(0, 1.12fr)
            minmax(300px, 0.88fr);

        gap: 38px;
    }

    .hero h1 {
        font-size:
            clamp(3.5rem, 6.6vw, 5.5rem);
    }

    .hero-media {
        min-height: 560px;
    }

    .hero-artwork {
        height:
            min(60vh, 560px);

        max-height: 560px;
        margin-right: -20px;
    }

    .expertise-grid {
        grid-template-columns:
            repeat(2, minmax(0, 1fr));
    }

    .expertise-grid article {
        border-bottom: 1px solid var(--border);
    }

    .contact-layout {
        grid-template-columns: 1fr;
        gap: 65px;
    }

    .contact-form,
    .contact-profiles {
        max-width: 900px;
    }

    .contact-profiles {
        display: grid;

        grid-template-columns:
            repeat(3, minmax(0, 1fr));

        column-gap: 30px;
    }

    .contact-profiles-label,
    .contact-privacy {
        grid-column: 1 / -1;
    }

}


/* =========================
   MOBILE NAV + TABLET STACKING
========================= */

@media (max-width: 900px) {

    .nav-container {
        height: 70px;
    }

    .menu-toggle {
        display: block;

        position: relative;
        z-index: 1200;
    }

    .site-nav {
        position: fixed;

        inset:
            70px
            0
            0
            0;

        width: 100vw;
        height: calc(100dvh - 70px);

        display: flex;
        flex-direction: column;
        align-items: stretch;

        gap: 0;

        padding:
            26px
            5%
            50px;

        overflow-y: auto;

        background: var(--background);

        transform: translateX(100%);

        transition: transform 220ms ease;
    }

    .site-nav.open {
        transform: translateX(0);
    }

    .site-nav a,
    .site-nav a:nth-child(6) {
        display: block;

        width: 100%;

        padding: 18px 0;

        border-bottom: 1px solid var(--border);

        color: var(--text-primary);

        font-size: 1.05rem;
    }

    .site-nav a::after {
        display: none;
    }

    .menu-toggle.active span:first-child {
        transform:
            translateY(3.7px)
            rotate(45deg);
    }

    .menu-toggle.active span:last-child {
        transform:
            translateY(-3.7px)
            rotate(-45deg);
    }

    .hero {
        min-height: auto;

        padding:
            78px
            0
            95px;
    }

    .hero-grid {
        grid-template-columns: 1fr;
        gap: 48px;
    }

    .hero-content {
        width: 100%;
    }

    .hero h1 {
        max-width: 760px;

        font-size:
            clamp(3.4rem, 10vw, 5.6rem);
    }

    .hero-statement {
        max-width: 700px;
    }

    .research-tags {
        max-width: 700px;
    }

    .hero-actions {
        width: 100%;
        max-width: 700px;
    }

    .hero-media {
        min-height: auto;
        width: 100%;

        padding: 5px 0 0;
    }

    .hero-artwork {
        width: auto;

        height:
            min(66vh, 560px);

        max-height: 560px;

        margin: 0 auto;
    }

    .section,
    .section-alt,
    .section-dark,
    .communication-section {
        padding: 95px 5%;
    }

    .section-grid,
    .about-grid,
    .communication-grid {
        grid-template-columns: 1fr;
        gap: 50px;
    }

    /* Keep Gene & Tonic compact on iPad/tablets.
       Feature images remain side-by-side and mosaic stays 4 columns. */

    .communication-feature-gallery,
    .communication-mosaic {
        max-width: 100%;
    }

    .communication-feature-gallery {
        grid-template-columns:
            repeat(2, minmax(0, 1fr));

        gap: 12px;
        margin-top: 52px;
    }

    .communication-mosaic {
        grid-template-columns:
            repeat(4, minmax(0, 1fr));

        gap: 8px;
        margin-top: 10px;
    }

    .section-header {
        display: block;
        margin-bottom: 50px;
    }

    .section-header > p {
        margin-top: 24px;
    }

    .about-portrait {
        width:
            min(100%, 420px);

        margin:
            38px
            auto
            0;
    }

    .research-grid {
        grid-template-columns: 1fr;
    }

    .timeline-item {
        grid-template-columns: 1fr;
        gap: 18px;
    }

    .publication-item {
        grid-template-columns: 1fr;
        gap: 25px;

        padding: 42px 0;
    }

    .publication-meta {
        flex-direction: row;
        align-items: center;
        flex-wrap: wrap;
    }

    .publications-footer {
        justify-content: stretch;
        margin-top: 36px;
    }

    .publication-record-link {
        width: 100%;
    }

    .expertise-grid {
        grid-template-columns: 1fr;
    }

    .expertise-grid article,
    .expertise-grid article:first-child,
    .expertise-grid article:last-child {
        padding: 30px 0;

        border-right: 0;
    }

    .activity-list div {
        grid-template-columns: 1fr;
        gap: 7px;
    }

    .contact-section {
        padding: 100px 5%;
    }

    .contact-layout {
        grid-template-columns: 1fr;
    }

    .contact-profiles {
        grid-template-columns: 1fr;
    }

    .contact-profiles-label,
    .contact-privacy {
        grid-column: 1;
    }

    .site-footer {
        min-height: auto;

        flex-direction: column;
        align-items: flex-start;
        justify-content: center;

        padding-top: 32px;
        padding-bottom: 32px;
    }

    .footer-links {
        justify-content: flex-start;

        gap:
            12px
            18px;
    }

}


/* =========================
   SMALL MOBILE
========================= */

@media (max-width: 600px) {

    .doctoral-credential {
        padding: 19px 20px;
        margin-bottom: 32px;
    }

    /* Gene & Tonic */

    .communication-feature-gallery {
        grid-template-columns: 1fr;

        width: 100%;
        max-width: 520px;

        gap: 10px;

        margin:
            42px
            auto
            0;
    }

    .communication-feature-photo {
        aspect-ratio: 4 / 3;
        border-radius: 14px;
    }

    .communication-mosaic {
        width: 100%;
        max-width: 520px;

        grid-template-columns:
            repeat(2, minmax(0, 1fr));

        gap: 7px;

        margin:
            8px
            auto
            0;
    }

    .communication-mosaic-item {
        aspect-ratio: 1 / 1;
        border-radius: 9px;
    }

    .communication-gallery-note {
        margin-top: 22px;

        font-size: 0.66rem;
        line-height: 1.7;
    }

    /* Phone hero */

    .hero {
        width: 90%;

        padding-top: 70px;
        padding-bottom: 80px;
    }

    .hero-grid {
        gap: 0;
    }

    .hero-media {
        display: none !important;
    }

    .eyebrow {
        font-size: 0.72rem;
        letter-spacing: 0.14em;
    }

    .hero h1 {
        margin-bottom: 28px;

        font-size:
            clamp(3.15rem, 14vw, 4.6rem);

        line-height: 0.94;
    }

    .hero-statement {
        margin-bottom: 30px;

        font-size:
            clamp(1.18rem, 5.8vw, 1.5rem);

        line-height: 1.42;
    }

    .research-tags {
        margin-bottom: 34px;
        gap: 8px;
    }

    .research-tags span {
        min-height: 36px;
        font-size: 0.75rem;
    }

    .hero-actions {
        grid-template-columns:
            minmax(0, 1fr)
            minmax(0, 1fr);

        gap: 10px;
    }

    .hero-actions .button {
        width: 100%;
        min-width: 0;

        padding: 0 12px;

        font-size: 0.82rem;
    }

    .section-heading h2,
    .section-header h2,
    .communication-grid h2 {
        font-size:
            clamp(2.5rem, 12vw, 4rem);
    }

    .research-feature {
        margin-top: 65px;
        padding-top: 42px;
    }

    .research-feature-header {
        margin-bottom: 30px;
    }

    .research-feature-header h3 {
        font-size:
            clamp(1.8rem, 8vw, 2.7rem);
    }

    .research-feature-header > p {
        margin-top: 20px;
        font-size: 0.92rem;
    }

    .research-feature-image {
        border-radius: 14px;
    }

    .research-feature figcaption {
        gap: 6px;
        margin-top: 14px;

        font-size: 0.72rem;
    }

    .form-row {
        grid-template-columns: 1fr;
        gap: 0;
    }

    .form-submit {
        width: 100%;
    }

}


/* =========================
   PHONE HERO — EXTRA SAFETY
========================= */

@media screen and (max-width: 600px) {

    .hero-media,
    .hero-artwork {
        display: none !important;
    }

    .hero-grid {
        gap: 0 !important;
    }

    .hero {
        min-height: auto;
        padding-bottom: 80px;
    }

}


/* =========================
   PHONE LANDSCAPE HERO
========================= */

@media (max-width: 950px) and (max-height: 700px) {

    .hero-media,
    .hero-artwork {
        display: none !important;
    }

    .hero-grid {
        grid-template-columns: 1fr !important;
        gap: 0 !important;
    }

    .hero {
        min-height: auto !important;

        padding-top: 55px;
        padding-bottom: 70px;
    }

    .hero-content {
        width: 100%;
    }

}


/* =========================
   REDUCED MOTION
========================= */

@media (prefers-reduced-motion: reduce) {

    html {
        scroll-behavior: auto;
    }

    * {
        transition: none !important;
    }

}